... and we are not talking about a beach on/near a reef... we are talking about a comparatively lifeless bed of sand in an aquarium with a staggeringly miniscule rate of water flow and exchange compared to the a given/analyzed portion of a beach.
In so many ways, we are talking about two different realities here. And so, we make concessions to the artifacts of captivity. That is the only biology I personally speak to/about. Aquarium science... not aquatic science, if you will
I do appreciate the references, Bugger... but 3/4 are 15-20 years old... the fourth is fast approaching that. Aquarium science and Aquatic science have evolved considerably since these excellent works were published, with all due respect my friend.
Anthony